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The master spy codenamed "Twilight" has spent his days on undercover missions, all for the dream of a better world. But one day, he receives a particularly difficult new order from command. For his mission, he must form a temporary family and start a new life?! A Spy/Action/Comedy about a one-of-a-kind family!

After rigorously training for three years, the ordinary Saitama has gained immense strength which allows him to take out anyone and anything with just one punch. He decides to put his new skill to good use by becoming a hero. However, he quickly becomes bored with easily defeating monsters, and wants someone to give him a challenge to bring back the spark of being a hero.

The demon-filled dimension known as "Mato" has opened to threaten humanity, and a type of "peach" tree that grows there yields fruit that confers special powers to those who eat it, giving humanity a fighting chance, but this only works for women. As such, as protectors of humanity, women rise in social status while men are relegated to second-class citizens scraping for basic occupation and acknowledgment.

In the American Old West, the world's greatest race is about to begin. Thousands line up in San Diego to travel over six thousand kilometers for a chance to win the grand prize of fifty million dollars. With the era of the horse reaching its end, contestants are allowed to use any kind of vehicle they wish. Competitors will have to endure grueling conditions, traveling up to a hundred kilometers a day through uncharted wastelands. The Steel Ball Run is truly a one-of-a-kind event. The youthful Johnny Joestar, a crippled former horse racer, has come to San Diego to watch the start of the race. There he encounters Gyro Zeppeli, a racer with two steel balls at his waist instead of a gun. Johnny witnesses Gyro using one of his steel balls to unleash a fantastical power, compelling a man to fire his gun at himself during a duel. In the midst of the action, Johnny happens to touch the steel ball and feels a power surging through his legs, allowing him to stand up for the first time in two years. Vowing to find the secret of the steel balls, Johnny decides to compete in the race, and so begins his bizarre adventure across America on the Steel Ball Run.